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known by the name of the "drum of the ear." The compages of bones consists of four, which are so disposed, and so hinge upon one another, as that if the membrane, the drum of the ear, vibrate, all the four are put in motion together; and, by the result of their action, work the base of that which is the last in the series, upon an aperture which it closes, and upon which it plays, and which aperture opens into the tortuous canals that lead to the brain. This last bone of the four is called the stapes. The office of the drum of the ear is to spread out an extended surface, capable of receiving the impressions of sound, and of being put by them into a state of vibration. The office of the stapes is to repeat these vibrations. It is a repeating frigate, stationed more within the line. From which account of its action may be understood, how the sensation of sound will be excited, by any thing which communicates a vibratory motion to the stapes, though not, as in all ordinary cases, through the intervention of the membrana tympani. This is done by solid bodies applied to the bones of the skull, as by a metal bar holden at one end between the teeth, and touching at the other end a tremulous body. It likewise appears to be done, in a considerable degree, by the air itself, even when this membrane, the drum of the ear, is greatly damaged. Either in the natural or preternatural state of the organ, the use of the chain of bones is to propagate the impulse in a direction towards the brain, and to propagate it with the advantage of a lever; which advantage consists in increasing the force and strength of the vibration, and at the same time diminishing the space through which it oscillates; both of which changes may augment or facilitate the still deeper action of the auditory nerves.

The benefit of the eustachian tube to the organ, may be made out upon known pneumatic principles. Behind the drum of the ear is a second cavity, or barrel, called the tympanum. The eustachian tube is a slender pipe, but sufficient for the passage of air, leading from this cavity into the back part of the mouth. Now, it would not have done to have had a vacuum in this cavity; for, in that case, the pressure of the atmosphere from without would have burst the membrane which covered it. Nor would it have done to have filled the cavity with lymph or any other secretion; which would necessarily have obstructed, both the vibration of the membrane, and the play of the small bones. Nor, lastly, would it have done to have occupied the space with confined air, because the expansion of that air by heat, or its contraction by cold, would have distended or relaxed the covering membrane, in a degree inconsistent with the purpose which it was assigned to execute. The only remaining expedient, and that for which the eustachian tube serves, is to open to this cavity a communication with the external air. In one word; it exactly answers the purpose of the hole in a drum.

The membrana tympani itself, likewise, deserves all the examination which can be made of it. It is not found in the ears of fish; which furnishes an additional proof of what indeed is indicated by every thing about it, that it is appropriated to the action of air, or of an elastic medium. It bears an obvious resemblance to the pelt or head of a drum, from which it takes its name. It resembles also a drum head in this principal property, that its use depends upon its tension. Tension is the state essential to it. Now we know, that in a drum, the pelt is carried over a hoop, and braced as occasion requires, by the means of strings attached to its circumference. In the membrane of the ear, the same purpose is provided for, more simply, but not less mechanically, nor less successfully, by a different expedient, viz. by the end of a bone (the handle of the malleus) pressing upon its centre. It is only in very large animals that the texture of this membrane can be discerned. In the Philosophical Transactions for the year 1800 (vol. i.), Mr. Everard Home has given some curious observations upon the ear, and the drum of the ear, of an elephant. He discovered in it, what he calls a radiated muscle, that is, straight muscular fibres, passing along the membrane from the circumference to the centre; from the bony rim which surrounds it towards the handle of the malleus to which the central part is attached. This muscle he supposes to be designed to bring the membrane into unison with different sounds; but then he also discovered, that this muscle itself cannot act, unless the membrane be drawn to a stretch, and kept in a due state of tightness, by what may be called a foreign force, viz. the action of the muscles of the malleus. Supposing his explanation of the use of the parts to

be just, our author is well founded in the reflection which he makes upon it: "that this mode of adapting the ear to different sounds, is one of the most beautiful applications of muscles in the body; the mechanism is so simple, and the variety of effects so great."

In another volume of the Transactions above referred to, and of the same year, two most curious cases are related, of persons who retained the sense of hearing, not in a perfect, but in a very considerable degree, notwithstanding the almost total loss of the membrane we have been describing. In one of these cases, the use here assigned to that membrane, of modifying the impressions of sound by change of tension, was attempted to be supplied by straining the muscles of the outward ear. "The external ear," we are told, "had acquired a distinct motion upward and backward, which was observable whenever the patient listened to any thing which he did not distinctly hear; when he was addressed in a whisper, the ear was seen immediately to move; when the tone of voice was louder, it then remained altogether motionless."

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It appears probable, from both these cases, that a collateral, if not principal, use of the membrane, is to cover and protect the barrel of the ear which lies behind it. Both the patients suffered from cold: one, a great increase of deafness from catching cold;" the other, "very considerable pain from exposure to a stream of cold air." Bad effects therefore followed from this cavity being left open to the external air; yet, had the Author of nature shut it up by any other cover than what was capable, by its texture, of receiving vibrations from sound, and, by its connexion with the interior parts, of transmitting those vibrations to the brain, the use of the organ, so far as can we judge, must have been entirely obstructed *.

Since the period at which Paley composed this work, many philosophical anatomists have engaged in researches relative to the structure of the ear. The chief of these are Weber, Malatides, Majendie, Monro, Lallemand, Deleau, Aland, Itard, Scarpa, and Saissy. Its delicate anatomy, its intricate mechanism, is consequently much more completely demonstrated than it was at the close of the eighteenth century. To enter into their details here, would be misplaced, and the general result is merely mentioned because delicacy and intricacy of workmanship proportionately evince the skill and the design required in the artificer.

It may be observed of the conch, or exterior part of the ear, that it is adapted excellently for the office it performs the collection of vibrations, or sound. All the art of the tyrant Dionysius, all the ingenuity stimulated into action by his gold, could not devise a better form to render audible the conversation of his state prisoners. He made their dungeon in the form of the human ear, two hundred and fifty feet in length, and eighty feet in height. The sounds within this were directed to a tympanum communicating with an adjoining room, where the regal monster crouched, and was enabled to distinguish even the whispers of those within his toils.

When a mechanician has invented an engine, a source of power, his ingenuity has not done all that is required of him; he has to adapt it to the purposes for which it is needed. The steam-engine, for instance, is made smaller, or increased in size proportionate to the power necessary; it has to be made, when employed for propelling carriages on a railway, of a form differing materially from that erected for the drainage of mines. Now this is all a proof of the mechanician's skill; it is a demonstration of a continuous design. The ear is a complete piece of mechanism; it is varied in all animals, in all is appropriate to their habits and wants; varies just as is required. Is it rational to say that the inference is not the same?

Birds, it is easy to perceive, would be impeded by an external ear; but its place is supplied by the feathers around the auditory orifice, which they can erect at

pleasure, so as to form a reflector. In some birds this is very remarkable. When passing rapidly through the air, the feathers lie close to the sides of the head.

Hares and other animals, whose safety depends upon flight, and whose perception of sound is most required to be directed behind them, have large ears naturally inclined backwards. Beasts of prey, and dogs, animals that are generally pursuing, seldom flying from a pursuer, have the conch of the ear directed forward; and those animals whose heads, as the ox when feeding, and the hound when in the chase, are held low, have pendulous ears, with the conch so turned as to be best adapted for gathering the sound that vibrates nearest the earth.

All animals that have to live partially or entirely under water, have their organs of hearing peculiarly adapted to their habits. The ear of the water shrew is closed against the entrance of water by a very curious valve.

The ear of the whale very closely resembles that of the human species. It, like them, passes the chief part of its life in the air; but when safety or hunger requires it to be submerged, the meatus, or opening of the ear, closes, and within the meatus is a small hard body more than an inch in length, attached by its small end to one side of the passage; this serves as a valve when the passage contracts.

Fish having no exterior ear, it was considered, until within comparatively a few years, that they were deficient in that sense to which it seemed necessary. Subsequent researches have shown the error of this conclusion. The necessity of the conch arises from terrestrial animals living in the air, which is a very imperfect conductor of sounds; an apparatus is required to collect a greater amount of vibrations to compensate for the deficiency of their intensity. To show that animals who live in the water, a denser and therefore better medium for conveying sounds, require no such exterior parts, let the reader, when diving, have a stone in each hand, and strike them together whilst beneath the water; he will be comparatively stunned by the concussion.

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CHAPTER IV.

ON THE SUCCESSION OF PLANTS AND ANIMALS.

THE generation of the animal no more accounts for the contrivance of the eye or ear, than, upon the supposition stated in a preceding chapter, the production of a watch, by the motion and mechanism of a former watch, would account for the skill and intention evidenced in the watch, so produced; than it would account for the disposition of the wheels, the catching of their teeth, the relation of the several parts of the works to one another, and to their common end; for the suitableness of their forms and places to their offices, for their connexion, their operation, and the useful result of that operation. I do insist most strenuously upon the correctness of this comparison; that it holds as to every mode of specific propagation; and that whatever was true of the watch, under the hypothesis above mentioned, is true of plants and animals.

I. To begin with the fructification of plants. Can it be doubted but that the seed contains a particular organisation? Whether a latent plantule with the means of temporary nutrition, or whatever else it be, it encloses an organisation suited to the germination of a new plant. Has the plant which produced the seed any thing more to do with that organisation, than the watch would have had to do with the structure of the watch which was produced in their course of its mechanical movement? I mean, Has it any thing at all to do with the contrivance? The maker and contriver of one watch, when he inserted within it a mechanism suited to the production of another watch, was, in truth, the maker and contriver of that other watch. All the properties of the new watch were to be referred to his agency: the design manifested in it, to his intention: the art, to him as the artist: the collocation of each part, to his placing: the action, effect, and use, to his counsel, intelligence, and workmanship. In producing it by the intervention of a former watch, he was only working by one set of tools instead of another. So it is with the plant, and the seed produced by it. Can any distinction be assigned between the two cases; between the producing watch, and the producing plant; both passive, unconscious substances; both, by the organisation which was given to them, producing their like, without understanding or design; both, that is, instruments?

II. From plants we may proceed to oviparous animals: from seed to eggs. Now I say, that the bird has the same concern in the formation of the egg which she lays, as the plant has in that of the seed which it drops: and no other, nor greater. The internal constitution of the egg is as much a secret to the hen, as if the hen were inanimate. Her will cannot alter it, or change a single feather of the chick. She can neither foresee nor determine of which sex her brood shall be, or how many of either: yet the thing produced shall be, from the first, very different in its make, according to the sex which it bears. So far, therefore, from adapting the means, she is not beforehand apprised of the effect. If there be concealed within that smooth shell a provision and a preparation for the production and nourishment of a new animal, they are not of her providing or preparing: if there be contrivance, it is none of hers. Although, therefore, there be the difference of life and perceptivity between the animal and the plant, it is a difference which enters not into the account. It is a foreign circumstance. It is a difference of properties not employed. The animal function and the vegetable function are alike destitute of any design which can operate upon the form of the thing produced. The plant has no design in producing the seed, no comprehension of the nature or use of what it produces: the bird with respect to its egg, is not above the plant with respect to its seed. Neither the one nor the other bears that sort of relation to what proceeds from them, which a joiner does to the chair which he makes. Now a cause, which bears this relation to the effect, is what we want, in order to account for the suitableness of means to an end, the fitness and fitting of one thing to another; and this cause the parent plant or animal does not supply.

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It is farther observable concerning the propagation of plants and animals, that the арраratus employed exhibits no resemblance to the thing produced in this respect holding an analogy with instruments and tools of art. The filaments, antheræ, and stigmata, of flowers, bear no more resemblance to the young plant, or even to the seed, which is formed by their intervention, than a chisel or a plane does to a table or chair. What then are the filaments, antheræ, and stigmata, of plants, but instruments strictly so called?*

III. We may advance from animals which bring forth eggs, to animals which bring forth their young alive: and of this latter class, from the lowest to the highest; from irrational to rational life, from brutes to the human species: without perceiving, as we proceed, any alteration whatever in the terms of the comparison. The rational animal does not produce its offspring with more certainty or success than the irrational animal; a man than a quadruped, a quadruped than a bird; nor (for we may follow the gradation through its whole scale) a bird than a plant; nor a plant than a watch, a piece of dead mechanism would do, upon the supposition which has already so often been repeated. Rationality, therefore, has nothing to do in the business. If an account must be given of the contrivance which we observe; if it be demanded, whence arose either the contrivance by which the young animal is produced, or the contrivance manifested in the young animal itself, it is not from the reason of the parent that any such account can be drawn. He is the cause of his offspring, in the same sense as that in which a gardener is the cause of the tulip which grows upon his parterre, and in no other. We admire the flower; we examine the plant; we perceive the conduciveness of many of its parts to their end and office: we observe a provision for its nourishment, growth, protection, and fecundity; but we never think of the gardener in all this. We attribute nothing of this to his agency; yet it may still be true, that without the gardener, we should not have had the tulip: just so it is with the succession of animals even of the highest order. For the contrivance discovered in the structure of the thing produced, we want a contriver. The parent is not that contriver. His consciousness decides that question. He is in total ignorance why that which is produced took its present form rather than any other. It is for him only to be astonished, by the effect. We can no more look therefore to the intelligence of the parent animal for what we are in search of, a cause of relation, and of subserviency of parts to their use, which relation and subserviency we see in the procreated body, than we can refer the internal conformation of an acorn to the intelligence of the oak from which it dropped, or the structure of the watch to the intelligence of the watch which produced it; there being no difference, as far as argument is concerned, between an intelligence which is not exerted, and an intelligence which does not exist.

CHAPTER V.

APPLICATION OF THE ARGUMENT CONTINued.

EVERY observation which was made in our first chapter, concerning the watch, may be repeated with strict propriety, concerning the eye; concerning animals; concerning plants; concerning, indeed, all the organised parts of the works of nature. As,

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matter, which she selects from old mortar, coal ashes, or other rubbish, with unvaried industry.

How essential this practice is to the formation of the shell, is shewn by the well ascertained fact, that when the hen is debarred access to calcareous matters, she lays her eggs without shells.

The hen then swallows, not by chance, but instinctively, two kinds of earth, flint-pebbles for the use of her gizzard, and chalk, for her eggshells. Both to us uninviting, tasteless substances, yet both essential to her health; and this is not the action of a single hen, but of all. Is not this an adaptation of the instinct of the bird to its wants?-Vauquelin, Ann. de Chim. vol. xxxiv. p. 71.

I. When we are inquiring simply after the existence of an intelligent Creator, imperfection, inaccuracy, liability to disorder, occasional irregularities may subsist in a considerable degree, without inducing any doubt into the question: just as a watch may frequently go wrong, seldom perhaps exactly right, may be faulty in some parts, defective in some, without the smallest ground of suspicion from thence arising that it was not a watch; not made; or not made for the purpose ascribed to it. When faults are pointed out, and when a question is started concerning the skill of the artist, or dexterity with which the work is executed, then indeed, in order to defend these qualities from accusation, we must be able, either to expose some intractableness and imperfection in the materials, or point out some invincible difficulty in the execution, into which imperfection and difficulty the matter of complaint. may be resolved; or, if we cannot do this, we must adduce such specimens of consummate art and contrivance proceeding from the same hand, as may convince the inquirer, of the existence, in the case before him, of impediments like those which we have mentioned, although, what from the nature of the case is very likely to happen, they be unknown and unperceived by him. This we must do in order to vindicate the artist's skill, or, at least, the perfection of it; as we must also judge of his intention, and of the provisions employed in fulfilling that intention, not from an instance in which they fail, but from the great plurality of instances in which they succeed. But, after all, these are different questions from the question of the artist's existence: or, which is the same, whether the thing before us be a work of art or not; and the questions ought always to be kept separate in the mind. So likewise it is in the works of nature. Irregularities and imperfections are of little or no weight in the consideration, when that consideration relates simply to the existence of a Creator. When the argument respects his attributes, they are of weight; but are then to be taken in conjunction (the attention is not to rest upon them, but they are to be taken in conjunction) with the unexceptionable evidences which we possess, of skill, power, and benevolence, displayed in other instances; which evidences may, in strength, number, and variety, be such, and may so overpower apparent blemishes, as to induce us, upon the most reasonable ground, to believe, that these last ought to be referred to some cause, though we be ignorant of it, other than defect of knowledge or of benevolence in the author.

II. There may be also parts of plants and animals, as there were supposed to be of the watch, of which, in some instances, the operation, in others, the use, is unknown. These form different cases; for the operation may be unknown, yet the use be certain. Thus it is with the lungs of animals. It does not, I think, appear that we are acquainted with the action of the air upon the blood, or in what manner that action is communicated by the lungs: yet we find that a very short suspension of their office destroys the life of the animal. In this case, therefore, we may be said to know the use, nay we experience the necessity of the organ, though we be ignorant of its operation. Nearly the same thing may be observed of what is called the lymphatic system. We suffer grievous inconveniences from its disorder, without being informed of the office which it sustains in the economy of our bodies. There may possibly also be some few examples of the second class, in which not only the operation is unknown, but in which experiments may seem to prove that the part is not necessary; or may leave a doubt, how far it is even useful to the plant or animal in which it is found. This is said to be the case with the spleen; which has been extracted from dogs, without any sensible injury to their vital functions. Instances of the former kind, namely, in which we cannot explain the operation, may be numerous; for they will be so in proportion to our ignorance. They will be more or fewer to different persons, and in different stages of science. Every improvement of knowledge diminishes their number. There is hardly perhaps a year passes, that does not, in the works of nature, bring some operation, or some mode of operation, to light, which was before undiscovered-probably unsuspected. Instances of the second kind, namely, where the part appears to be totally useless, I believe to be extremely rare; compared with the number of those, of which the use is evident, they are beneath any assignable proportion; and, perhaps, have never been submitted to a trial and examination sufficiently accurate, long enough continued, or often enough repeated. No accounts which I have seen are satisfactory. The mutilated animal may live and grow fat, (as was the case of the dog deprived of its spleen), yet may be defective in some other of its functions;

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