The same remark applies to the views of Professor Geddes on the laws of growth which have determined certain essential features in the morphology of plants and animals. The attempt to substitute these laws for those of variation and natural selection has failed in cases where we can apply a definite test, as in that of the origin of spines on trees and shrubs; while the extreme diversity of vegetable structure and form among the plants of the same country and of the same natural order, of itself affords a proof of the preponderating influence of variation and natural selection in keeping the many diverse forms in harmony with the highly complex and ever-changing environment.

Lastly, we have seen that Professor Weismann's theory of the continuity of the germ-plasm and the consequent nonheredity of acquired characters, while in perfect harmony with all the well-ascertained facts of heredity and development, adds greatly to the importance of natural selection as the one invariable and ever-present factor in all organic change, and that which can alone have produced the temporary fixity combined with the secular modification of species. While admitting, as Darwin always admitted, the co-operation of the fundamental laws of growth and variation, of correlation and heredity, in determining the direction of lines of variation or in the initiation of peculiar organs, we find that variation and natural selection are ever-present agencies, which take possession, as it were, of every minute change originated by these fundamental causes, check or favour their further development, or modify them in countless varied ways according to the varying needs of the organism. Whatever other causes have been at work, Natural Selection is supreme, to an extent which even Darwin himself hesitated to claim for it. The more we study it the more we are convinced of its overpowering importance, and the more confidently we claim, in Darwin's own words, that it "has been the most important, but not the exclusive, means of modification."

CHAPTER XV

DARWINISM APPLIED TO MAN

General identity of human and animal structure—Rudiments and variations showing relation of man to other mammals-The embryonic development of man and other mammalia-Diseases common to man and the lower animals-The animals most nearly allied to man— The brains of man and apes-External differences of man and apes— Summary of the animal characteristics of man-The geological antiquity of man-The probable birthplace of man-The origin of the moral and intellectual nature of man-The argument from continuity-The origin of the mathematical faculty-The origin of the musical and artistic faculties-Independent proof that these faculties have not been developed by natural selection-The interpretation of the facts-Concluding remarks.

OUR review of modern Darwinism might fitly have terminated with the preceding chapter; but the immense interest that attaches to the origin of the human race, and the amount of misconception which prevails regarding the essential teachings of Darwin's theory on this question, as well as regarding my own special views upon it, induce me to devote a final chapter to its discussion.

To any one who considers the structure of man's body, even in the most superficial manner, it must be evident that it is the body of an animal, differing greatly, it is true, from the bodies of all other animals, but agreeing with them in all essential features. The bony structure of man classes him as a vertebrate; the mode of suckling his young classes him as a mammal; his blood, his muscles, and his nerves, the structure of his heart with its veins and arteries, his lungs and his whole respiratory and circulatory systems, all closely correspond to those of other mammals, and are often almost identical with

them. He possesses the same number of limbs terminating in the same number of digits as belong fundamentally to the mammalian class. His senses are identical with theirs, and his organs of sense are the same in number and occupy the same relative position. Every detail of structure which is common to the mammalia as a class is found also in man, while he only differs from them in such ways and degrees as the various species or groups of mammals differ from each other. If, then, we have good reason to believe that every existing group of mammalia has descended from some common ancestral form—as we saw to be so completely demonstrated in the case of the horse tribe,—and that each family, each order, and even the whole class must similarly have descended from some much more ancient and more generalised type, it would be in the highest degree improbable—so improbable as to be almost inconceivable—that man, agreeing with them so closely in every detail of his structure, should have had some quite distinct mode of origin. Let us, then, see what other evidence bears upon the question, and whether it is sufficient to convert the probability of his animal origin into a practical certainty.

Rudiments and Variations as Indicating the Relation of Man to other Mammals.

All the higher animals present rudiments of organs which, though useless to them, are useful in some allied group, and are believed to have descended from a common ancestor in which they were useful. Thus there are in ruminants rudiments of incisor teeth which, in some species, never cut through the gums; many lizards have external rudimentary legs; while many birds, as the Apteryx, have quite rudimentary wings. Now man possesses similar rudiments, sometimes constantly, sometimes only occasionally present, which serve intimately to connect his bodily structure with that of the lower animals. Many animals, for example, have a special muscle for moving or twitching the skin. In man there are remnants of this in certain parts of the body, especially in the forehead, enabling us to raise our eyebrows; but some persons have it in other parts. A few persons are able to move the whole scalp so as to throw off any object placed on the head,

and this property has been proved, in one case, to be inherited. In the outer fold of the ear there is sometimes a projecting point, corresponding in position to the pointed ear of many animals, and believed to be a rudiment of it. In the alimentary canal there is a rudiment-the vermiform appendage of the cæcumwhich is not only useless, but is sometimes a cause of disease and death in man; yet in many vegetable feeding animals it is very long, and even in the orang-utan it is of considerable length and convoluted. So, man possesses rudimentary bones of a tail concealed beneath the skin, and, in some rare cases, this forms a minute external tail.

The variability of every part of man's structure is very great, and many of these variations tend to approximate towards the structure of other animals. The courses of the arteries are eminently variable, so that for surgical purposes it has been necessary to determine the probable proportion of each variation. The muscles are so variable that in fifty cases the muscles of the foot were found to be not strictly alike in any two, and in some the deviations were considerable; while in thirty-six subjects Mr. J. Wood observed no fewer than 558 muscular variations. The same author states that in a single male subject there were no fewer than seven muscular variations, all of which plainly represented muscles proper to various kinds of apes. The muscles of the hands and arms-parts which are so eminently characteristic of man-are extremely liable to vary, so as to resemble the corresponding muscles of the lower animals. That such variations are due to reversion to a former state of existence Mr. Darwin thinks highly probable, and he adds: "It is quite incredible that a man should, through mere accident, abnormally resemble certain apes in no less than seven of his muscles, if there had been no genetic connection between them. On the other hand, if man is descended from some ape-like creature, no valid reason can be assigned why certain muscles should not suddenly reappear after an interval of many thousand generations, in the same manner as, with horses, asses, and mules, dark coloured stripes suddenly reappear on the legs and shoulders, after an interval of hundreds, or inore probably of thousands of generations."

"1

1 Descent of Man, pp. 41-43; also pp. 13-15.

The Embryonic Development of Man and other Mammalia.

The progressive development of any vertebrate from the ovum or minute embryonic egg affords one of the most marvellous chapters in Natural History. We see the contents of the ovum undergoing numerous definite changes, its interior dividing and subdividing till it consists of a mass of cells, then a groove appears marking out the median line or vertebral column of the future animal, and thereafter are slowly developed the various essential organs of the body. After describing in some detail what takes place in the case of the ovum of the dog, Professor Huxley continues: "The history of the development of any other vertebrate animal, lizard, snake, frog, or fish tells the same story. There is always to begin with, an egg having the same essential structure as that of the dog; the yelk of that egg undergoes division or segmentation, as it is called, the ultimate products of that segmentation constitute the building materials for the body of the young animal; and this is built up round a primitive groove, in the floor of which a notochord is developed. Furthermore, there is a period in which the young of all these animals resemble one another, not merely in outward form, but in all essentials of structure, so closely, that the differences between them are inconsiderable, while in their subsequent course they diverge more and more widely from one another. And it is a general law that the more closely any animals resemble one another in adult structure, the larger and the more intimately do their embryos resemble one another; so that, for example, the embryos of a snake and of a lizard remain like one another longer than do those of a snake and a bird; and the embryos of a dog and of a cat remain like one another for a far longer period than do those of a dog and a bird, or of a dog and an opossum, or even than those of a dog and a monkey."1

We thus see that the study of development affords a test of affinity in animals that are externally very much unlike each other; and we naturally ask how this applies to man. Is he developed in a different way from other mammals, as we should certainly expect if he has had a distinct and

1 Man's Place in Nature, p. 64.