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In the first place, they practically agree in upsetting, each in its own way, the generally-received definition of species, and in sweeping away the ground of their objective existence in Nature. The orthodox conception of species is that of lineal descent: all the descendants of a common parent, and no other, constitute a species; they have a certain identity because of their descent, by which they are supposed to be recognizable. So naturalists had a distinct idea of what they meant by the term species, and a practical rule, which was hardly the less useful because difficult to apply in many cases, and because its application was indirect: that is, the community of origin had to be inferred from the likeness; such degree of similarity, and such only, being held to be conspecific as could be shown or reasonably inferred to be compatible with a common origin. And the usual concurrence of the whole body of naturalists (having the same data before them) as to what forms are species attests the value of the rule, and also indicates some real foundation for it in Nature. But if species were created in numberless individuals over broad spaces of territory, these individuals are connected only in idea, and species differ from varieties on the one hand, and from genera, tribes, etc., on the other, only in degree; and no obvious natural reason remains for fixing upon this or that degree as specific, at least no natural standard, by which the opinions of different naturalists may be correlated. Species upon this view are enduring, but subjective and ideal. Any three or more of the human races, for example, are species or not species, according to the bent of the naturalist's mind. Darwin's

theory brings us the other way to the same result. In his view, not only all the individuals of a species are descendants of a common parent, but of all the related species also. Affinity, relationship, all the terms which naturalists use figuratively to express an underived, unexplained resemblance among species, have a literal meaning upon Darwin's system, which they little sus. pected, namely, that of inheritance. Varieties are the latest offshoots of the genealogical tree in "an unlineal" order; species, those of an earlier date, but of no definite distinction; genera, more ancient species, and so on. The human races, upon this view, likewise may or may not be species according to the notions of each naturalist as to what differences are specific; but, if not species already, those races that last long enough are sure to become so. It is only a question of time.

How well the simile of a genealogical tree illustrates the main ideas of Darwin's theory the following extract from the summary of the fourth chapter shows:

"It is a truly wonderful fact-the wonder of which we are apt to overlook from familiarity—that all animals and all plants throughout all time and space should be related to each other in group subordinate to group, in the manner which we everywhere behold-namely, varieties of the same species most closely related together, species of the same genus less closely and unequally related together, forming sections and sub-genera, species of distinct genera much less closely related, and genera related in different degrees, forming sub-families, families, orders, sub-classes, and classes. The several subordinate groups in any class cannot be ranked in a single file, but seem rather to be clustered round points, and these round other points, and so on in almost endless cycles. On the view that each species has been independently created, I can see no explanation of this

tains the essence of the whole, yet much of the aroma escapes in the treble distillation, or is so concentrated that the flavor is lost to the general or even to the scientific reader. The volume itself-the proof-spirit -is just condensed enough for its purpose. It will be far more widely read, and perhaps will make deeper impression, than the elaborate work might have done, with all its full details of the facts upon which the author's sweeping conclusions have been grounded. At least it is a more readable book: but all the facts that can be mustered in favor of the theory are still likely to be needed.

Who, upon a single perusal, shall pass judgment upon a work like this, to which twenty of the best years of the life of a most able naturalist have been devoted? And who among those naturalists who hold a position that entitles them to pronounce summarily upon the subject, can be expected to divest himself for the nonce of the influence of received and favorite systems? In fact, the controversy now opened is not likely to be settled in an off-hand way, nor is it desirable that it should be. A spirited conflict among opinions of every grade must ensue, which— to borrow an illustration from the doctrine of the book before us-may be likened to the conflict in Nature among races in the struggle for life, which Mr. Darwin describes; through which the views most favored by facts will be developed and tested by "Natural Selection," the weaker ones be destroyed in the process, and the strongest in the long-run alone survive.

The duty of reviewing this volume in the American Journal of Science would naturally devolve upon

the principal editor, whose wide observation and profound knowledge of various departments of natural history, as well as of geology, particularly qualify him. for the task. But he has been obliged to lay aside. his pen, and to seek in distant lands the entire repose from scientific labor so essential to the restoration of his health-a consummation devoutly to be wished, and confidently to be expected. Interested as Mr. Dana would be in this volume, he could not be expected to accept its doctrine. Views so idealistic as those upon which his "Thoughts upon Species"1 are grounded, will not harmonize readily with a doctrine so thoroughly naturalistic as that of Mr. Darwin. Though it is just possible that one who regards the kinds of elementary matter, such as oxygen and hydrogen, and the definite compounds of these elementary matters, and their compounds again, in the mineral kingdom, as constituting species, in the same sense, fundamentally, as that of animal and vegetable species, might admit an evolution of one species from another in the latter as well as the former case.

Between the doctrines of this volume and those of the other great naturalist whose name adorns the titlepage of this journal [Mr. Agassiz], the widest divergence appears. It is interesting to contrast the two, and, indeed, is necessary to our purpose; for this contrast brings out most prominently, and sets in strongest light and shade, the main features of the theory of the origination of species by means of Natural Selection. The ordinary and generally-received view assumes the independent, specific creation of each kind of plant 1 Article in this Journal, vol. xxiv., p. 305.

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DARWINIANA.

and animal in a primitive stock, which reproduces its
like from generation to generation, and so continues
the species. Taking the idea of species from this
perennial succession of essentially similar individuals,
the chain is logically traceable back to a local origin in
a single stock, a single pair, or a single individual,
from which all the individuals composing the species
have proceeded by natural generation. Although the
similarity of progeny to parent is fundamental in the
conception of species, yet the likeness is by no means
absolute; all species vary more or less, and some vary
remarkably-partly from the influence of altered cir-
cumstances, and partly (and more really) from un-
known constitutional causes which altered conditions
favor rather than originate. But these variations are
supposed to be mere oscillations from a normal state,
and in Nature to be limited if not transitory; so that
the primordial differences between species and species
at their beginning have not been effaced, nor largely
obscured, by blending through variation. Conse-
quently, whenever two reputed species are found to
blend in Nature through a series of intermediate forms,
community of origin is inferred, and all the forms,
however diverse, are held to belong to one species.
Moreover, since bisexuality is the rule in Nature
(which is practically carried out, in the long-run, far
more generally than has been suspected), and the
heritable qualities of two distinct individuals are min-
gled in the offspring, it is supposed that the general

1 "Species tot sunt, quot diversas formas ab initio produxit Infini-
tum Ens; quæ formæ, secundum generationis inditas leges, produxere
plures, at sibi semper similes."-Linn. Phil. Bot., 99, 157.

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