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1833.] ZOOLOGY OF NORTH AND SOUTH AMERICA.

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ruminant, discovered by MM. Lund and Clausen in the caves of Brazil, are highly interesting facts with respect to the geographical distribution of animals. At the present time, if we divide America, not by the Isthmus of Panama, but by the southern part of Mexico* in lat. 20°, where the great table-land presents an obstacle to the migration of species, by affecting the climate, and by forming, with the exception of some valleys and of a fringe of low land on the coast, a broad barrier; we shall then have the two zoological provinces of North and South America strongly contrasted with each other. Some few species alone have passed the barrier, and may be considered as wanderers from the south, such as the puma, opossum, kinkajou, and peccari. South America is characterized by possessing many peculiar gnawers, a family of monkeys, the llama, peccari, tapir, opossums, and, especially, several genera of Edentata, the order which includes the sloths, ant-eaters, and armadillos. North America, on the other hand, is characterized (putting on one side a few wandering species) by numerous peculiar gnawers, and by four genera (the ox, sheep, goat, and antelope) of hollow-horned ruminants, of which great division South America is not known to possess a single species. Formerly, but within the period when most of the now existing shells were living, North America possessed, besides hollow-horned ruminants, the elephant, mastodon, horse, and three genera of Edentata, namely, the Megatherium, Megalonyx, and Mylodon. Within nearly this same periods (as proved by the shells at Bahia Blanca) South America possessed, as we have just seen, a mastodon, horse, hollow-horned ruminant, and the same three genera (as well as several others) of the Edentata. Hence it is evident that North and South America, in having within a late geological period these several genera in common, were much more closely related in the character of their terrestrial inhabitants than they now are.

*This is the geographical division followed by Lichtenstein, Swainson, Erichson, and Richardson. The section from Vera Cruz to Acapulco, given by Humboldt in the Polit. Essay on Kingdom of N. Spain, will show how immense a barrier the Mexican table-land forms. Dr. Richardson, in his admirable Report on the Zoology of N. America read before the Brit. Assoc. 1836 (p. 157), talking of the identification of a Mexican animal with the Synetheres prehensilis, says, "We do not know with what propriety, but if correct, it is, if not a solitary instance, at least very nearly so, of a rodent animal being common to North and South America.'

The more I reflect on this case, the more interesting it appears : I know of no other instance where we can almost mark the period and manner of the splitting up of one great region into two well-characterized zoological provinces. The geologist, who is fully impressed with the vast oscillations of level which have affected the earth's crust within late periods, will not fear to speculate on the recent elevation of the Mexican platform, or, more probably, on the recent submergence of land in the West Indian Archipelago, as the cause of the present zoological separation of North and South America. The South American character of the West Indian mammals * seems to indicate that this archipelago was formerly united to the southern continent, and that it has subsequently been an area of subsidence.

When America, and especially North America, possessed its elephants, mastodons, horse, and hollow-horned ruminants, it was much more closely related in its zoological characters to the temperate parts of Europe and Asia than it now is. As the remains of these genera are found on both sides of Behring's Straits and on the plains of Siberia, we are led to look to the north-western side of North America as the former point of communication between the Old and so-called New World. And as so many species, both living and extinct, of these same genera inhabit and have inhabited the Old World, it seems most probable that the North American elephants, mastodons, horse, and hollowhorned ruminants migrated, on land since submerged near Behring's Straits, from Siberia into North America, and thence, on land since submerged in the West Indies, into South America, where for a time they mingled with the forms characteristic of that southern continent, and have since become extinct.

While travelling through the country, I received several vivid descriptions of the effects of a late great drought; and the account of this may throw some light on the cases where vast

*See Dr. Richardson's Report, p. 157; also L'Institut, 1837, p. 253. Cuvier says the kinkajou is found in the larger Antilles, but this is doubtful. M. Gervais states that the Didelphis crancrivora is found there. It is certain that the West Indies possess some mammifers peculiar to themselves. A tooth of a mastodon has been brought from Bahama: Edin. New Phil. Journ. 1826, p. 395.

See the admirable Appendix by Dr. Buckland to Beechey's Voyage, also the writings of Chamisso in Kotzebue's Voyage.

1833.]

THE GREAT DROUGHT.

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numbers of animals of all kinds have been embedded together. The period included between the years 1827 and 1830 is called the "gran seco," or the great drought. During this time so little rain fell, that the vegetation, even to the thistles, failed; the brooks were dried up, and the whole country assumed the appearance of a dusty high road. This was especially the case in the northern part of the province of Buenos Ayres and the southern part of St. Fé. Very great numbers of birds, wild animals, cattle, and horses perished from the want of food and water. A man told me that the deer* used to come into his courtyard to the well, which he had been obliged to dig to supply his own family with water; and that the partridges had hardly strength to fly away when pursued. The lowest estimation of the loss of cattle in the province of Buenos Ayres alone, was taken at one million head. A proprietor at San Pedro had previously to these years 20,000 cattle; at the end not one remained. San Pedro is situated in the middle of the finest country; and even now abounds again with animals; yet, during the latter part of the "gran seco," live cattle were brought in vessels for the consumption of the inhabitants. The animals roamed from their estancias, and, wandering far southward, were mingled together in such multitudes, that a government commission was sent from Buenos Ayres to settle the disputes of the owners. Sir Woodbine Parish informed me of another and very curious source of dispute; the ground being so long dry, such quantities of dust were blown about, that in this open country the landmarks became obliterated, and people could not tell the limits of their estates.

I was informed by an eyewitness that the cattle in herds of thousands rushed into the Parana, and being exhausted by hunger they were unable to crawl up the muddy banks, and thus were

* In Capt. Owen's Surveying Voyage (vol. ii. p. 274) there is a curious account of the effects of a drought on the elephants, at Benguela (west coast of Africa). "A number of these animals had some time since entered the town, in a body, to possess themselves of the wells, not being able to procure any water in the country. The inhabitants mustered, when a desperate conflict ensued, which terminated in the ultimate discomfiture of the invaders, but not until they had killed one man, and wounded several others." The town is said to have a population of nearly three thousand! Dr. Malcolmson informs me, that during a great drought in India the wild animals entered the tents of some troops at Ellore, and that a hare drank out of a vessel held by the adjutant of the regiment.

drowned. The arm of the river which runs by San Pedro was so full of putrid carcasses, that the master of a vessel told me that the smell rendered it quite impassable. Without doubt several hundred thousand animals thus perished in the river: their bodies when putrid were seen floating down the stream; and many in all probability were deposited in the estuary of the Plata. All the small rivers became highly saline, and this caused the death of vast numbers in particular spots; for when an animal drinks of such water it does not recover. Azara describes* the fury of the wild horses on a similar occasion, rushing into the marshes, those which arrived first being overwhelmed and crushed by those which followed. He adds that more than once he has seen the carcasses of upwards of a thousand wild horses thus destroyed. I noticed that the smaller streams in the Pampas were paved with a breccia of bones, but this probably is the effect of a gradual increase, rather than of the destruction at any one period. Subsequently to the drought of 1827 to '32, a very rainy season followed, which caused great floods. Hence it is almost certain that some thousands of the skeletons were buried by the deposits of the very next year. What would be the opinion of a geologist, viewing such an enormous collection of bones, of all kinds of animals and of all ages, thus embedded in one thick earthy mass? Would he not attribute it to a flood having swept over the surface of the land, rather than to the common order of things?†

October 12th.-I had intended to push my excursion further, but not being quite well, I was compelled to return by a balandra, or one-masted vessel of about a hundred tons' burden, which was bound to Buenos Ayres. As the weather was not fair, we moored early in the day to a branch of a tree on one of the islands. The Parana is full of islands, which undergo a constant round of decay and renovation. In the memory of the master several large ones had disappeared, and others again had been formed and protected by vegetation. They are composed of muddy sand, without even the smallest pebble, and were then * Travels, vol. i. p. 374.

+ These droughts to a certain degree seem to be almost periodical; I was told the dates of several others, and the intervals were about fifteen years.

1833.]

HABITS OF THE JAGUAR,

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about four feet above the level of the river; but during the periodical floods they are inundated. They all present one character; numerous willows and a few other trees are bound together by a great variety of creeping plants, thus forming a thick jungle. These thickets afford a retreat for capybaras and jaguars. The fear of the latter animal quite destroyed all pleasure in scrambling through the woods. This evening I had not proceeded a hundred yards, before finding indubitable signs of the recent presence of the tiger, I was obliged to come back. On every island there were tracks; and as on the former excursion "el rastro de los Indios" had been the subject of conversation, so in this was "el rastro del tigre."

The wooded banks of the great rivers appear to be the favourite haunts of the jaguar; but south of the Plata, I was told that they frequented the reeds bordering lakes: wherever they are, they seem to require water. Their common prey is the capybara, so that it is generally said, where capybaras are numerous there is little danger from the jaguar. Falconer states that near the southern side of the mouth of the Plata there are many jaguars, and that they chiefly live on fish; this account I have heard repeated. On the Parana they have killed many wood-cutters, and have even entered vessels at night. There is a man now living in thẹ Bajada, who, coming up from below when it was dark, was seized on the deck; he escaped, however, with the loss of the use of one arm. When the floods drive these animals from the islands, they are most dangerous. I was told that a few years since a very large one found its way into a church at St. Fé: two padres entering one after the other were killed, and a third, who came to see what was the matter, escaped with difficulty. The beast was destroyed by being shot from a corner of the building which was unroofed. They commit also at these times great ravages among cattle and horses. It is said that they kill their prey by breaking their necks. If driven from the carcass, they seldom return to it. The Gauchos say that the jaguar, when wandering about at night, is much tormented by the foxes yelping as they follow him. This is a curious coincidence with the fact which is generally affirmed of the jackals accompanying, in a similarly officious manner; the East Indian tiger. The jaguar is a noisy animal, roaring much by night, and especially before bad weather.

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