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FIG.

20. RECOGNITION OF EDICNEMUS VERMICULATUS AND E. SENEGALENSIS (from Seebohm's Charadriada)

PAGE

223

21. RECOGNITION OF CURSORIUS CHALCOPTERUS AND C. GALLICUS (from Seebohm's Charadriado)

224

22. RECOGNITION OF SCOLOPAX MEGALA AND S. STENURA (from Seebohm's Charadriada)

225

23. METHONA PSIDII AND LEPTALIS ORISE.

241

24. OPTHALMIS LINCEA AND ARTAXA SIMULANS (from the Official Narrative of the Voyage of the Challenger)

247

25. WINGS OF ITUNA ILIONE AND THYRIDIA MEGISTO (from Proceedings of the Entomological Society).

251

26. MYGNIMIA AVICULUS AND COLOBORHOMBUS FASCIATIPENNIS 27. MIMICKING INSECTS FROM THE PHILIPPINES (from Semper's Animal Life).

259

260

28. MALVA SYLVESTRIS AND M. ROTUNDIFOLIA (from Lubbock's British Wild Flowers in Relation to Insects)

311

29. LYTHRUM SALICARIA, THREE FORMS OF (from Lubbock's British Wild Flowers in Relation to Insects)

312

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30. ORCHIS PYRAMIDALIS (from Darwin's Fertilisation of Orchids). 314 31. HUMMING-BIRD FERTILISING MARCGRAVIA NEPENTHOIDES 320

32. DIAGRAM OF MEAN HEIGHT OF LAND AND DEPTH OF OCEANS 345 33. GEOLOGICAL DEVELOPMENT OF THE HORSE TRIBE (from Huxley's American Addresses)

388

34. DIAGRAM ILLUSTRATING THE GEOLOGICAL DISTRIBUTION OF PLANTS (from Ward's Sketch of Palæobotany).

402

35. TRANSFORMATION OF ARTEMIA SALINA TO A. MILHAUSENII (from Semper's Animal Life) .

426

36. BRANCHIPUS STAGNALIS AND ARTEMIA SALINA (from Semper's

Animal Life).

427

37. CHIMPANZEE (TROGLODYTES NIGER)

454

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CHAPTER I

WHAT ARE SPECIES," AND WHAT IS MEANT BY

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Definition of species-Special creation-The early Transmutationists—
Scientific opinion before Darwin-The problem before Darwin—
The change of opinion effected by Darwin-The Darwinian theory
-Proposed mode of treatment of the subject.

THE title of Mr. Darwin's great work is-On the Origin of
Species by means of Natural Selection and the Preservation of
Favoured Races in the Struggle for Life. In order to ap-
preciate fully the aim and object of this work, and the
change which it has effected not only in natural history but
in many other sciences, it is necessary to form a clear con-
ception of the meaning of the term "species," to know what
was the general belief regarding them at the time when Mr.
Darwin's book first appeared, and to understand what he
meant, and what was generally meant, by discovering their
origin." It is for want of this preliminary knowledge that
the majority of educated persons who are not naturalists are
so ready to accept the innumerable objections, criticisms, and
difficulties of its opponents as proofs that the Darwinian
theory is unsound, while it also renders them unable to ap-
preciate, or even to comprehend, the vast change which that
theory has effected in the whole mass of thought and opinion
on the great question of evolution.

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The term "species" was thus defined by the celebrated botanist De Candolle: "A species is a collection of all the individuals which resemble each other more than they resemble anything else, which can by mutual fecundation

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produce fertile individuals, and which reproduce themselves by generation, in such a manner that we may from analogy suppose them all to have sprung from one single individual." And the zoologist Swainson gives a somewhat similar definition: "A species, in the usual acceptation of the term, is an animal which, in a state of nature, is distinguished by certain peculiarities of form, size, colour, or other circumstances, from another animal. It propagates, 'after its kind,' individuals perfectly resembling the parent; its peculiarities, therefore, are permanent." 1

To illustrate these definitions we will take two common English birds, the rook (Corvus frugilegus) and the crow (Corvus corone). These are distinct species, because, in the first place, they always differ from each other in certain slight peculiarities of structure, form, and habits, and, in the second place, because rooks always produce rooks, and crows produce crows, and they do not interbreed. It was therefore concluded that all the rooks in the world had descended from a single pair of rooks, and the crows in like manner from a single pair of crows, while it was considered impossible that crows could have descended from rooks or vice versâ. The

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origin" of the first pair of each kind was a mystery. Similar remarks may be applied to our two common plants, the sweet violet (Viola odorata) and the dog violet (Viola canina). These also produce their like and never produce each other or intermingle, and they were therefore each supposed to have sprung from a single individual whose origin" was unknown. But besides the crow and the rook there are about thirty other kinds of birds in various parts of the world, all so much like our species that they receive the common name of crows; and some of them differ less from each other than does our crow from our rook. These are all species of the genus Corvus, and were therefore believed to have been always as distinct as they are now, neither more nor less, and to have each descended from one pair of ancestral crows of the same identical species, which themselves had an unknown "origin." Of violets there are more than a hundred different kinds in various parts of the world, all differing very slightly from each other and forming distinct. 1 Geography and Classification of Animals, p. 350.

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