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virginianus are registered, we see a remarkable spreading out of the spots, showing in some of the characters a tendency to segregation into two or more groups of individuals, each varying considerably from the mean.

In order fully to appreciate the teaching of these diagrams,

VARIATION OF

ICTERUS BALTIMORE. 20.3

Tail.

Wing.

Tarsus.

Middle Toe.

Hind Toe.

Bill, Length.

Bill, Width.

FIG. 9.

we must remember, that, whatever kind and amount of variations are exhibited by the few specimens here compared, would be greatly extended and brought into symmetrical form if large numbers-thousands or millions—were subjected to the same process of measurement and registration. We know, from the general law which governs variations from a mean value, that with increasing numbers the range.

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of variation of each part would increase also, at first rather rapidly and then more slowly; while gaps and irregularities

Curves of Variation

FIG. 11.

would be gradually filled up, and at length the distribution of the dots would indicate a tolerably regular curve of double curvature like those shown in Fig. 11. The great divergence

of the dots, when even a few specimens are compared, shows that the curve, with high numbers, would be a flat one like the lower curve in the illustration here given. This being the case it would follow that a very large proportion of the total number of individuals constituting a species would diverge considerably from its average condition as regards each part or organ; and as we know from the previous diagrams of variation (Figs. 1 to 7) that each part varies to a considerable extent, independently, the materials constantly ready for natural selection CARDINALIS VIRGINIANUS. 58 specimens. Florida.

Tail.

Length of Bird.

Wing.

(From Allen's Birds of Florida. p.281)

FIG. 12

to act upon are abundant in quantity and very varied in kind. Almost any combination of variations of distinct parts will be available, where required; and this, as we shall see further on, obviates one of the most weighty objections which have been urged against the efficiency of natural selection in producing new species, genera, and higher groups.

Variation in the Mammalia.

Owing to the generally large size of this class of animals, and the comparatively small number of naturalists who study them, large series of specimens are only occasionally examined

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and compared, and thus the materials for determining the question of their variability in a state of nature are comparatively scanty. The fact that our domestic animals belonging to this group, especially dogs, present extreme varieties not surpassed even by pigeons and poultry among birds, renders it almost certain that an equal amount of variability exists in the wild state; and this is confirmed by the example of a species of squirrel (Sciurus carolinensis), of which sixteen specimens, all males and all taken in Florida, were measured and tabulated by Mr. Allen. The diagram here given shows, that, both the general amount of the variation and the independent variability of the several members of the body, accord completely with the variations so common in the class of birds; while their amount and their independence of each other are even greater than usual.

Variation in the Internal Organs of Animals.

In case it should be objected that the cases of variation hitherto adduced are in the external parts only, and that there is no proof that the internal organs vary in the same manner, it will be advisable to show that such varieties also occur. It is, however, impossible to adduce the same amount of evidence in this class of variation, because the great labour of dissecting large numbers of specimens of the same species. is rarely undertaken, and we have to trust to the chance observations of anatomists recorded in their regular course of study.

It must, however, be noted that a very large proportion of the variations already recorded in the external parts of animals necessarily imply corresponding internal variations. When feet and legs vary in size, it is because the bones vary; when the head, body, limbs, and tail change their proportions, the bony skeleton must also change; and even when the wing or tail feathers of birds become longer or more numerous, there is sure to be a corresponding change in the bones which support and the muscles which move them. I will, however, give a few cases of variations which have been directly observed.

Mr. Frank E. Beddard has kindly communicated to me some remarkable variations he has observed in the internal

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