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HOPLOPHONEUS MENTALIS sp. nov.

Type No. 12515, Princeton University Geological Museum, collecting locality 1015A1, left ramus of the lower jaw with the canine and third and fourth premolars in place (Fig. 1), secured by the 1920 South Dakota expedition from the uppermost levels of the Titanotherium beds (Chadron formation), two to two and one half feet below the thin local bed of white limestone at the contact between the Chadron and Brule formations (Oreodon beds), and two and a half to three feet above titanothere bones in place, in the valley of Indian Creek, near Taylor's ranch, west of Hart Table in Pennington County (locality shown in Fig. 2 of Plate 1 of the preceding paper, in about the center of the picture).

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So far as I am aware, this is the first Hoplophoneus to be described from the Titanotherium beds and is strikingly differentiated from all of the species of the overlying Oreodon beds by the extraordinary depth of the chin (as suggested in the specific name proposed), a character reminiscent of the great development of this structure in Hatcher's Eusmilus dakotensis from the Protoceras beds, to which the new form seems to be transitional.

1 Williston's measurements, Kansas Univ. Quarterly, Vol. III., No. 3, p. 72, 1895.

2 Specimens in the Princeton Geological Museum used by G. I. Adams, in defining these species.

From the table of measurements it will readily be seen that Hoplophoneus mentalis is considerably smaller than the large H. occidentalis, approximating in size individuals of the insolens and robustus groups. A comparison with the specimens in the Princeton collection used by Adams in defining these two forms shows that, although the jaw depth back of the flange is the same in H. robustus and H. insolens as in the new form, the flange is absolutely larger in the latter and tapers less rapidly in width distally; both premolars are smaller, and in p4 the paraconid is disproportionately smaller, than in H. insolens (Princeton Univ. Geol. Mus. No. 11372) and is out of line with the other two cusps, toward the inner side of the jaw as in Eusmilus and hoplophonids in general, and has sharper cutting edges than in H. insolens; also the alveolus for mi is very much wider transversely, in front, than in the latter species. Between H. robustus and H. mentalis there is the same striking difference in the shape of the jaw flange, which is decidedly V-shaped in robustus, while in the new species it is U-shaped and of the Eusmilus type. The third premolar is as small as in robustus, but rakes backward a bit stronger and has a larger posterior cuspule. To the paraconid of p4 the same remarks are applicable as in the comparison with H. insolens.

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